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Professor Joe Cain

UCL Professor of History and Philosophy of Biology

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  • [ 2026-08-11 ] Comparing Tempo and Mode (1944) with Major Features (1953) by George Gaylord Simpson Synthesis Period
  • [ 2026-08-11 ] Major Features of Evolution (1953) by George Gaylord Simpson Key Texts
  • [ 2026-08-11 ] Variation and Evolution in Plants (1950) by G. Ledyard Stebbins Key Texts
  • [ 2026-08-11 ] Tempo and Mode in Evolution (1944) by George Gaylord Simpson Key Texts
  • [ 2026-08-11 ] Systematics and the Origin of Species (1942) by Ernst Mayr Key Texts
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Key Texts

Key Texts in the Synthesis Period of Evolutionary Studies

Excerpt from photograph of conference attendees at 1947 Princeton conference organised by National Research Council's Committee on Common Problems of Genetics, Paleontology, and Systematics
Excerpt from photograph of conference attendees at 1947 Princeton conference organised by National Research Council’s Committee on Common Problems of Genetics, Paleontology, and Systematics.

The synthesis period of evolutionary studies, clustering in the late 1930s and 1940s, saw new approaches and new interdisciplinary conversations driving a spirit of innovation and transformation. Advocates celebrated a “modern synthesis” and focused attention on books in the Columbia Biological Series as textbooks for their common enterprise. Books in this series included:

  • Theodosius Dobzhansky’s (1937) Genetics and the Origin of Species
  • Ernst Mayr’s (1942) Systematics and the Origin of Species
  • George Gaylord Simpson’s (1944) Tempo and Mode in Evolution, which he later replaced with (1953) Major Features of Evolution
  • G. Ledyard Stebbins’s (1950) Variation and Evolution in Plants

There was much more to the history of the synthesis period than books in the Columbia Biological Series and their authors, together labelled as the “New York Circle”. This is commonly forgotten by those seeking to describe this period. At the same time, these key texts create a cluster that merits detailed analysis.

George Gaylord Simpson’s (1953) The Major Features of Evolution. Title page
Key Texts

Major Features of Evolution (1953) by George Gaylord Simpson

George Gaylord Simpson‘s (1902-1984) book, Major Features of Evolution, was published in 1953 by Columbia University Press in their Columbia Biological Series. This book played an important part in the expansion of evolutionary studies in […]
G. Ledyard Stebbins’s (1950) Variation and Evolution in Plants. Title page
Key Texts

Variation and Evolution in Plants (1950) by G. Ledyard Stebbins

G. Ledyard Stebbins‘ (1906-2000) book, Variation and Evolution in Plants, was published in 1950 by Columbia University Press in their Columbia Biological Series. This book played an important part in the expansion of evolutionary studies […]
George Gaylord Simpson’s (1944) Tempo and Mode in Evolution. Title page
Key Texts

Tempo and Mode in Evolution (1944) by George Gaylord Simpson

George Gaylord Simpson‘s (1902-1984) book, Tempo and Mode in Evolution, was published in 1944 by Columbia University Press in their Columbia Biological Series. This book played an important part in the expansion of evolutionary studies […]
Ernst Mayr’s (1942) Systematics and the Origin of Species from the Viewpoint of a Zoologist. Title page
Key Texts

Systematics and the Origin of Species (1942) by Ernst Mayr

Ernst Mayr‘s (1904-2005) book, Systematics and the Origin of Species from the Viewpoint of a Zoologist was published in 1942 by Columbia University Press in their Columbia Biological Series. This book played an important part […]
Theodosius Dobzhansky’s (1937) Genetics and the Origin of Species. Title page
Key Texts

Genetics and the Origin of Species (1937) by Theodosius Dobzhansky

Theodosius Dobzhansky‘s (1900-1975) book, Genetics and the Origin of Species, was first published in 1937 by Columbia University Press in their Columbia Biological Series. This book played an important part in the expansion of evolutionary […]

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Recent posts

  • Comparing Tempo and Mode (1944) with Major Features (1953) by George Gaylord Simpson
  • Major Features of Evolution (1953) by George Gaylord Simpson
  • Variation and Evolution in Plants (1950) by G. Ledyard Stebbins
  • Tempo and Mode in Evolution (1944) by George Gaylord Simpson
  • Systematics and the Origin of Species (1942) by Ernst Mayr

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Professor Joe Cain 2026